Research Publications

2026

  • Chopra, M., Rowlands, G., Stevens, G.M., Fernando, D., T, M., Laglbauer, B.J., Karnad, D. and Davis, K.J., 2026. Fewer devil rays in the sea: Evidence of declining mobulid populations off India's coast. Conservation Science and Practice, p.e70352.

  • Gonzalez-Pestana, A., Kyne, P.M., García-Rodríguez, E., Charles, R., Brito, V.B., Armstrong, A.O.,…Fernando, D.,…Neranjan, A.,…Tanna, A.,… & Jabado R.W., 2026. Critical habitats for sharks and rays in Asia remain largely unprotected. Biodiversity and Conservation, 35(7), p.179.

  • Laglbauer, B.J., D'Costa, N.G., Stewart, J.D., Palacios, M.D., Cronin, M., Fernando, D., Lezama-Ochoa, N., Armstrong, A.O., Jabado, R.W., Fowler, S. and Lawson, J.M., 2026. Global manta and devil ray population declines: Closing policy and management gaps to reduce fisheries mortality. Biological Conservation, 313, p.111589.

2025

  • Cardeñosa, D., Babcock, E.A., Shea, S.K., Zhang, H., Feldheim, K.A., Yang, F., Gale, S.W., Fernando, D., Tanna, A., Warwick, L. and D. Chapman, D., 2025. International trade regulations take a limited bite out of the shark fin trade. Science Advances, 11(45), p.eadz2821.

  • Feutry, P., Foster, S., Grewe, P.M., Aulich, J., Lansdell, M., Clear, N., Cooper, S., Williams, A., Johnson, G., Dilrukshi, T. and Wudianto, 2025. Genome scans reveal extensive population structure in three neritic tuna and tuna-like species in the Indian Ocean. ICES Journal of Marine Science, 82(2), p.fsae162.

  • Kipperman, M.J., Jabado, R.W., Haque, A.B., Fernando, D., Anjani, P.A.D.L., Spaet, J.L. and Humble, E., 2025. Population structure and genetic diversity of the Critically Endangered bowmouth guitarfish (Rhina ancylostomus) in the Northwest Indian Ocean. Peer Community Journal, 5.

  • Hanselman, K.M., Gobiraj, R. and Jensen, K., 2025. Revision of the Lecanicephalidean Genus Flapocephalus Deshmukh, 1979 (Eucestoda) from Cowtail Rays (Genus Pastinachus Rüppell) (Myliobatiformes: Dasyatidae Jordan and Gilbert) with the Description of Four New Species. The Journal of Parasitology, 111(5), pp.582-605.

    Text is available at: https://doi.org/10.1645/24-92

  • Palacios, M.D., Weiand, L., Laglbauer, B.J., Cronin, M.R., Fowler, S., Jabado, R.W., Ko Gyi, T., Fernando, D., De Bruyne, G., Shea, S.K. and Hilton, P., 2025. Global assessment of manta and devil ray gill plate and meat trade: conservation implications and opportunities. Environmental Biology of Fishes, 108(4), pp.611-638.

  • Kashey, S., Doumbouya, F., Laglbauer, B., Ender, I., Fernando, D., Stevens, G.M. and Simpfendorfer, C.A., 2025. Guinean landing surveys of Bentfin devil rays add insight into threatened elasmobranch catch. Environmental Biology of Fishes, pp.1-17.

  • Bond, M.E., Booth, H., Tanna, A., Fowler, S.L., Polo-Silva, C.J., Shea, K.H.S., Cardiec, F., Mansur, E.F., Jabado, R.W. (2025). Trade regulations drive improved global shark and ray management. Marine Policy, 180, Article 106733.

    Text is available at: https://doi.org/10.1016/j.marpol.2025.106733-02386-9

2024

  • Mohanraj, T., Rajathy, T.J., Stevens, G., Fernando, D. and Shunmugavel, C., 2024. Length-weight Relationship of the Endangered Devil Ray Mobula mobular (Bonnaterre, 1778) off Gulf of Mannar, India: Length-weight Relationship of Mobula mobular off Gulf of Mannar, India. Fishery Technology, 61(3).

  • Goetze, J.S., Heithaus, M.R., MacNeil, M.A., Harvey, E., Simpfendorfer, C.A., Heupel,…Fernando, D. … & Chapman, D.D., 2024. Directed conservation of the world’s reef sharks and rays. Nature Ecology & Evolution, pp.1-11.

    Text is available at: https://www.nature.com/articles/s41559-024-02386-9

  • Finucci, B., Pacoureau, N., Rigby, C.L., Matsushiba, J.H., Faure-Beaulieu, N., Sherman, C.S., … Fernando, D. … & Dulvy, N.K., 2024. Fishing for oil and meat drives irreversible defaunation of deepwater sharks and rays. Science, 383(6687), pp.1135-1141.

    Text is available at: https://doi.org/10.1126/science.ade9121

2023

  • Postaire, B. D., Devloo-Delva, F., Brunnschweiler, J. M., Charvet, P., Chen, X., Cliff, G., ... Fernando, D. … Pathirana, B.M. … & Mona, S., 2023. Global genetic diversity and historical demography of the Bull Shark. Journal of Biogeography, NA.

    Text is available at: https://doi.org/10.1111/jbi.14774

  • Humble, E., Hosegood, J., Carvalho, G., de Bruyn, M., Creer, S., Stevens, G. M., ... Fernando, D. … & Ogden, R., 2023. Comparative population genomics of manta rays has global implications for management. bioRxiv, 2023-06.

  • Simpfendorfer C.A, Heithaus M., Heupel M., MacNeil A.M., Meekan M., Harvey, M., ... Bown, R.M.K. ... Fernando, D. ... Perera, N. ... Tanna, A. ... and Chapman D.D., 2023. Widespread diversity deficits of coral reef sharks and rays. Science 380, 1155–1160. https://doi.org/10.1038/s41586-020-2519-y

  • Kyne, P.M., di Sciara, G.N., Morera, A.B., Charles, R., Rodríguez, E.G., Fernando, D., Pestana, A.G., Priest, M. and Jabado, R.W., 2023. Important Shark and Ray Areas: a new tool to optimize spatial planning for sharks. Oryx, 57(2), pp.146-147.

  • Nikolic, N., Devloo‐Delva, F., Bailleul, D., Noskova, E., Rougeux, C., Delord, C., Borsa, P., Liautard‐Haag, C., Hassan, M., Marie, A.D. Feutry, P., …, Fernando, D., … Arnaud-Haond, S., 2023. Stepping up to genome scan allows stock differentiation in the worldwide distributed blue shark Prionace glauca. Molecular Ecology, 32(5), pp.1000-1019.

  • Devloo‐Delva, F., Burridge, C.P., Kyne, P.M., Brunnschweiler, J.M., Chapman, D.D., Charvet, P., Chen, X., Cliff, G., Daly, R., Drymon, J.M. Espinoza, M., Fernando, D., … Feutry, P., 2023. From rivers to ocean basins: The role of ocean barriers and philopatry in the genetic structuring of a cosmopolitan coastal predator. Ecology and Evolution, 13(2), p.e9837.

  • Sherman, C.S., Simpfendorfer, C.A., Pacoureau, N., Matsushiba, J.H., Yan, H.F., Walls, R.H., Rigby, C.L., VanderWright, W.J., Jabado, R.W., Pollom, R.A., … Fernando D., … Dulvy, N.K., 2023. Half a century of rising extinction risk of coral reef sharks and rays. Nature Communications, 14(1), p.15.

2022

  • Perera, N., Lokupitiya, E., Halwatura, D. and Udagedara, S., 2022. Quantification of blue carbon in tropical salt marshes and their role in climate change mitigation. Science of The Total Environment, p.153313.

  • Jones, B.L.H., Unsworth, R.K.F., Nordlund, L.M., Ambo-Rappe, R., Nafie, Y.A.L., Lopez, M.R., Udagedara, S., Cullen-Unsworth, L.C., 2022. Local ecological knowledge reveals change in seagrass social–ecological systems. Oceans 202, 3(3), 419-430.

  • Jones, B.L.H., Unsworth, R.K.F., Nordlund, L.M., Eklof, J.S., Ambo-Rappe, R., Carly, F., Jiddawi, N.S., Nafie, Y.A.L., Udagedara, S., Cullen-Unsworth, L.C., 2022. Dependence on seagrass fisheries governed by household income and adaptive capacity. Ocean and Coastal Management, 225, 106247.

  • Gorman, D., Jayakody., Kodikara, S., Udagedara, S., Dahanayaka, D.D.G.L., Kithsiri, B., Ranawana, M. P., Kumara, Pahalawattaarachchi. V., Thavanayagam, M., Steven, A., Jagath, A.A.L., Gunawardana, Vanderklift, M.A., 2022. Past, present and future of blue carbon ecosystems in Sri Lanka: conservation, restoration and policy. Oceanography and Marine Biology: An Annual Review Volume 60.

2021

  • Fernando, D., and Stewart, J.D., 2021. High bycatch rates of manta and devil rays in the “small-scale” artisanal fisheries of Sri Lanka. PeerJ. https://doi.org/10.7717/peerj.11994

    Infographics (by Shanika Perera):‍ ‍ENGLISH‍ ‍SINHALA‍ ‍TAMIL‍ ‍

  • Dulvy, N.K., Pacoureau, N., Rigby, C.L., Pollom, R.A., Jabado, R.W., Ebert, D.A., Finucci, B., Pollock, C.M., Cheok, J., Derrick, D.H., Herman, K.B., Sherman, S., VanderWright, W.J., Lawson, J.M., Walls, R.H.L., Carlson, J.K., Charvet, P., Bineesh, K.K., Fernando, D., Ralph, G.M., Matsushiba, J.H., Hilton-Taylor, C., Fordham, S.V., Simpfendorfer, C.A., 2021. Overfishing drives over one-third of all sharks and rays toward a global extinction crisis. Current Biology. https://doi.org/10.1016/j.cub.2021.08.062

  • Tanna, A., Fernando, D., Gobiraj, R., Pathirana, B.M., Thilakaratna, S., and Jabado, R.W., 2021. Where have all the sawfishes gone? Perspectives on declines of these Critically Endangered species in Sri Lanka. Aquatic Conservation: Marine and Freshwater Ecosystems.https://doi.org/10.1002/aqc.3617

    Infographics (by Shanika Perera):‍ ‍ENGLISH‍ ‍SINHALA‍ ‍TAMIL‍ ‍

  • Caira, J.N., Otto, K., Fernando, D., and Jensen, K., 2021. Three new species of “tetraphyllidean” cestodes from an undescribed bamboo shark (Orectolobiformes: Hemiscylliidae) in Sri Lanka. Folia Parasitologica 68. https://doi.org/10.14411/fp.2021.004

  • Pacoureau, N., Rigby, C.L., Kyne, P.M., Sherley, R.B., Winker, H., Carlson, J.K., Fordham, S. v., Barreto, R., Fernando, D., Francis, M.P., Jabado, R.W., Herman, K.B., Liu, K.-M., Marshall, A.D., Pollom, R.A., Romanov, E. v., Simpfendorfer, C.A., Yin, J.S., Kindsvater, H.K., and Dulvy, N.K., 2021. Half a century of global decline in oceanic sharks and rays. Nature 589, 567–571. https://doi.org/10.1038/s41586-020-03173-9

2020

  • Hosegood, J., Humble, E., Ogden, R., de Bruyn, M., Creer, S., Stevens, G., Abudaya, M., Bassos-Hull, K., Bonfil, R., Fernando, D. Foote, A.D., Hipperson, H., Jabado, R.W., Kaden, J., Moazzam, M., Peel, L.R., Pollett, S., Ponzo, A., Poortvliet, M., Salah, J., Senn, H., Stewart, J.D., Wintner, S., and Carvalho, G., 2019. Phylogenomics and species delimitation for effective conservation of manta and devil rays. Molecular Ecology, 29: 4783– 4796. https://doi.org/10.1111/mec.15683

  • MacNeil A.M., Chapman D., Heupel M., Simpfendorfer C.A, Heithaus M., Meekan M., ... Bown, R.M.K. ... Fernando, D. ... Perera, N. ... Tanna, A. ... and Cinner J.E., 2020. Global status and conservation potential of reef sharks. Nature 583, 801–806. https://doi.org/10.1038/s41586-020-2519-y

  • Notarbartolo di Sciara, G., Stevens, G. and Fernando, D., 2020. The giant devil ray Mobula mobular (Bonnaterre, 1788) is not giant, but it is the only spinetail devil ray. Marine Biodiversity Records 13, 4. https://doi.org/10.1186/s41200-020-00187-0

2019

  • Notarbartolo di Sciara, G., Adnet, S., Bennett, M., Broadhurst, M.K., Fernando, D., Jabado, R.W., Laglbauer, B.J. and Stevens, G., 2019. Taxonomic status, biological notes, and conservation of the longhorned pygmy devil ray Mobula eregoodoo (Cantor, 1849). Aquatic Conservation: Marine and Freshwater Ecosystems.

  • Hosegood, J., Humble, E., Ogden, R., de Bruyn, M., Creer, S., Stevens, G., Abudaya, M., Bassos-Hull, K., Bonfil, R., Fernando, D. Foote, A.D., Hipperson, H., Jabado, R.W., Kaden, J., Moazzam, M., Peel, L.R., Pollett, S., Ponzo, A., Poortvliet, M., Salah, J., Senn, H., Stewart, J.D., Wintner, S., and Carvalho, G., 2019. Genome-wide data for effective conservation of manta and devil ray species. bioRxiv, p.458141.

    Preprint is available at: https://www.biorxiv.org/content/10.1101/458141v2

  • Fernando, D., Bown, R.M., Tanna, A., Gobiraj, R., Ralicki, H., Jockusch, E.L., Ebert, D.A., Jensen, K. and Caira, J.N., 2019. New insights into the identities of the elasmobranch fauna of Sri Lanka. Zootaxa, 4585(2), pp.201-238.

2018

  • Stevens, G., Fernando, D. and Di Sciara, G.N., 2018. Guide to the Manta and Devil Rays of the World. Princeton University Press.

  • Stewart, J.D., Jaine, F.R., Armstrong, A.J., Armstrong, A.O., Bennett, M.B., Burgess, K.B., Couturier, L.I., Croll, D.A., Cronin, M.R., Deakos, M.H., Dudgeon, C.L., Fernando, D., Froman, N., Germanov, E.S., Hall, M.A., Hinojosa-Alvarez, S., Hosegood, J.E., Kashiwagi, T., Laglbauer, B.J.L., Lezama-Ochoa, N., Marshall, A.D., McGregor, F., Notarbartolo di Sciara, G., Palacios, M.D., Peel, L.R., Richardson, A.J., Rubin, R.D., Townsend, K.A., Venables, S.K., Stevens, G.M.W., 2018. Research priorities to support effective manta and devil ray conservation. Frontiers in Marine Science, 5, 2018;314.

  • Jabado, R.W., Kyne, P.M., Pollom, R.A., Ebert, D.A., Simpfendorfer, C.A., Ralph, G.M., Al Dhaheri, S.S., Akhilesh, K.V., Ali, K., Ali, M.H., Al Mamari, T.M., Bineesh, K.K., El Hassan, I.S., Fernando, D., Grandcourt, E.M., Khan, M.M., Moore, A.B.M., Owfi, F., Robinson, D.P., Romanov, E., Soares, A., Spaet, J.L.Y., Tesfamichael, D., Valinassab, T., Dulvy, N.K. 2018. Troubled waters: Threats and extinction risk of the sharks, rays and chimaeras of the Arabian Sea and adjacent waters. Fish and Fisheries, 19(6), pp.1043-1062.

  • Jones, B.L., Unsworth, R.K., Udagedara, S. and Cullen-Unsworth, L.C., 2018. Conservation concerns of small-scale fisheries: by-catch impacts of a shrimp and finfish fishery in a Sri Lankan Lagoon. Frontiers in Marine Science, 5, p.52.

2017

  • Perera, N., Dahanayaka, D.D.G.L. and Udagedara, S., 2017. Habitat Preference and Population Structure of Two Data Deficient Seahorse (Syngnathidae) Species. OUSL Journal, 12(2), pp.75-97.

  • Udagedara, S., Fernando, D., Perera, N., Tanna, A. and Bown, R., 2017. A first record of Halodule pinifolia Miki den Hartog, and new locality of nationally endangered Halophila beccarii Asch, from the eastern coast of Sri Lanka. International Journal of Aquatic Biology, 5(5), pp.328-335.

  • Stewart, J.D., Rohner, C.A., Araujo, G., Avila, J., Fernando, D., Forsberg, K., Ponzo, A., Rambahiniarison, J.M., Kurle, C.M. and Semmens, B.X., 2017. Trophic overlap in mobulid rays: insights from stable isotope analysis. Marine Ecology Progress Series, 580, pp.131-151.

  • Abudaya, M., Ulman, A., Salah, J., Fernando, D., Wor, C. and di Sciara, G.N., 2018. Speak of the devil ray (Mobula mobular) fishery in Gaza. Reviews in fish biology and fisheries, 28(1), pp.229-239.

  • Lawson, J.M., Fordham, S.V., O’Malley, M.P., Davidson, L.N., Walls, R.H., Heupel, M.R., Stevens, G., Fernando, D., Budziak, A., Simpfendorfer, C.A., Ender, I., Francis, M.P., Notarbartolo di Sciara, G., and Dulvy, N.K., 2017. Sympathy for the devil: a conservation strategy for devil and manta rays. PeerJ, 5, p.e3027.

  • Kularatne, H. and Udagedara, S., 2017. First record of the Woolly-necked Stork Ciconia episcopus Boddaert, 1783 (Aves: Ciconiiformes: Ciconiidae) breeding in the lowland wet zone of Sri Lanka. Journal of Threatened Taxa, 9(1), pp.9738-9742.

2016

  • Stewart, J.D., Beale, C.S., Fernando, D., Sianipar, A.B., Burton, R.S., Semmens, B.X. and Aburto-Oropeza, O., 2016. Spatial ecology and conservation of Manta birostris in the Indo-Pacific. Biological Conservation, 200, pp.178-183.

  • Camargo, S.M., Coelho, R., Chapman, D., Howey-Jordan, L., Brooks, E.J., Fernando, D., Mendes, N.J., Hazin, F.H., Oliveira, C., Santos, M.N. and Foresti, F., 2016. Structure and genetic variability of the oceanic whitetip shark, Carcharhinus longimanus, determined using mitochondrial DNA. PloS one, 11(5), p.e0155623.

  • Notarbartolo di Sciara, G., Fernando, D., Adnet, S., Cappetta, H. and Jabado, R.W., 2017. Devil rays (Chondrichthyes: Mobula) of the Arabian Seas, with a redescription of Mobula kuhlii (Valenciennes in Müller and Henle, 1841). Aquatic Conservation: Marine and Freshwater Ecosystems, 27(1), pp.197-218.

2015

  • Croll, D.A., Dewar, H., Dulvy, N.K., Fernando, D., Francis, M.P., Galván‐Magaña, F., Hall, M., Heinrichs, S., Marshall, A., Mccauley, D. and Newton, K.M., 2016. Vulnerabilities and fisheries impacts: the uncertain future of manta and devil rays. Aquatic Conservation: Marine and Freshwater Ecosystems, 26(3), pp.562-575.

  • Fernando, D., Perera, N. and Ebert, D.A., 2015. First record of the megamouth shark, Megachasma pelagios,(Chondrichthyes: Lamniformes: Megachasmidae) from Sri Lanka, northern Indian Ocean. Marine Biodiversity Records, 8. p.e75.

  • Ooi, M.S., Townsend, K.A., Bennett, M.B., Richardson, A.J., Fernando, D., Villa, C.A. and Gaus, C., 2015. Levels of arsenic, cadmium, lead and mercury in the branchial plate and muscle tissue of mobulid rays. Marine pollution bulletin, 94(1-2), pp.251-259.

  • Fernando, D., 2014. Range Extension of the Bull Shark Carcharhinus leucas (Carcharhiniformes: Carcharhinidae) to include Sri Lanka. Journal of the Wildlife and Nature Protection Society of Sri Lanka (Loris), 27(1), pp.26-27.

2014

  • Morris, J.P., Thatje, S., Ravaux, J., Shillito, B., Fernando, D. and Hauton, C., 2014. Acute combined pressure and temperature exposures on a shallow-water crustacean: novel insights into the stress response and high pressure neurological syndrome. Comparative Biochemistry and Physiology Part A: Molecular & Integrative Physiology, 181, pp.9-17.

  • Poortvliet, M., Olsen, J.L., Croll, D.A., Bernardi, G., Newton, K., Kollias, S., O’Sullivan, J., Fernando, D., Stevens, G., Magaña, F.G. and Seret, B., 2014. A dated molecular phylogeny of manta and devil rays (Mobulidae) based on mitogenome and nuclear sequences. Molecular Phylogenetics and Evolution, 83, pp.72-85.